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- Page: i
- First Published: 13 October 2021

Cover: Holotype of Archaeocidaris ivanovi from the Carboniferous of Russia (PIN 5348/192). Specimen (including matrix) approx. 87 mm wide. (https://doi.org/10.1002/spp2.1280)
Issue Information
Original Articles
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A previously missing link in the evolution of dasytine soft-winged flower beetles from Cretaceous Charentese amber (Coleoptera, Melyridae)
- Pages: 1753-1764
- First Published: 11 April 2021
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Phylogenetic affinities and morphology of the Pliocene cathartiform Dryornis pampeanus Moreno & Mercerat
- Pages: 1765-1780
- First Published: 16 April 2021
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A new species of the Cambrian bivalved euarthropod Pectocaris with axially differentiated enditic armatures
- Pages: 1781-1792
- First Published: 28 April 2021
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Silurian (late Llandovery – Wenlock) conodont fauna and biostratigraphy from the Yanbian area of Sichuan Province, south-west China
- Pages: 1793-1824
- First Published: 28 April 2021
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Morphological–phylogenetic analysis of the late Cenozoic Chlamydini von Teppner (Bivalvia, Pectinidae) of southern South America
- Pages: 1825-1846
- First Published: 30 April 2021
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Rosselichnidae ifam. nov.: burrows with concentric, spiral or eccentric lamination
- Pages: 1847-1875
- First Published: 08 June 2021
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A Burgess Shale mandibulate arthropod with a pygidium: a case of convergent evolution
- Pages: 1877-1894
- First Published: 15 June 2021
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Evolutionary pattern of Metacaremys gen. nov. (Rodentia, Octodontidae) and its biochronological implications for the late Miocene and early Pliocene of southern South America
- Pages: 1895-1917
- First Published: 15 June 2021
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First pan-trionychid turtle (Testudines, Pan-Trionychidae) from the Palaeogene of Africa
- Pages: 1919-1926
- First Published: 16 June 2021
Open Access
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Late Cretaceous hydrothermal vent communities from the Troodos ophiolite, Cyprus: systematics and evolutionary significance
- Pages: 1927-1947
- First Published: 22 June 2021
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A revision of Vernicomacanthus Miles with comments on the characters of stem-group chondrichthyans
- Pages: 1949-1976
- First Published: 29 June 2021
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Panderodus from the Waukesha Lagerstätte of Wisconsin, USA: a primitive macrophagous vertebrate predator
- Pages: 1977-1993
- First Published: 09 July 2021
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Morphological disparity in the evolution of the ophthalmosaurid forefin: new clues from the Upper Jurassic of Argentina
- Pages: 1995-2020
- First Published: 14 July 2021
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An unpredicted ancient colonization of the West Indies by North American rodents: dental evidence of a geomorph from the early Oligocene of Puerto Rico
- Pages: 2021-2039
- First Published: 15 July 2021
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A Silurian ophiuroid with soft-tissue preservation
- Pages: 2041-2047
- First Published: 18 July 2021
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An early Eocene fossil from the British London Clay elucidates the evolutionary history of the enigmatic Archaeotrogonidae (Aves, Strisores)
- Pages: 2049-2064
- First Published: 18 July 2021
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Earliest fossil record of the genus Tetradium (Rutaceae) in Asia: implications for its evolution and palaeoecology
- Pages: 2065-2074
- First Published: 18 July 2021
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A new unenlagiine (Theropoda, Dromaeosauridae) from the Upper Cretaceous of Brazil
- Pages: 2075-2099
- First Published: 05 August 2021
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Attenborough's beauty: exceptional pattern preservation in a frog-legged leaf beetle from the Eocene Green River Formation, Colorado (Coleoptera, Chrysomelidae, Sagrinae)
- Pages: 2101-2112
- First Published: 05 August 2021
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Evolutionary–phylogenetic pathway of the Cretaceous ammonite genus Aegocrioceras and its relationship to Juddiceras spp. and Crioceratites spp.
- Pages: 2113-2139
- First Published: 15 August 2021
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New and revised small shelly fossil record from the lower Cambrian of northern Iran
- Pages: 2141-2181
- First Published: 19 August 2021
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Palynology of a short sequence of the Lower Devonian Beartooth Butte Formation at Cottonwood Canyon (Wyoming): age, depositional environments and plant diversity
- Pages: 2183-2204
- First Published: 20 August 2021
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Phylogeny, disparity and mass extinction response in the trilobite order Harpetida
- Pages: 2205-2225
- First Published: 22 August 2021
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Lophotrochozoa in the Cambrian evolutionary radiation and the Pelagiella problem
- Pages: 2227-2244
- First Published: 25 August 2021
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The first Silurian trilobite with three-dimensionally preserved soft parts reveals novel appendage morphology
- Pages: 2245-2253
- First Published: 25 August 2021
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New information on the Jurassic lepidosauromorph Marmoretta oxoniensis
- Pages: 2255-2278
- First Published: 01 September 2021
Discussion
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Comment on: The ichnogenus Lumbricaria Münster from the Upper Jurassic of Germany interpreted as faecal strings of ammonites
- Pages: 2279-2281
- First Published: 10 June 2021
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Response to Comment on: The ichnogenus Lumbricaria Münster from the Upper Jurassic of Germany interpreted as faecal strings of ammonites
- Pages: 2283-2284
- First Published: 11 June 2021