Revision of the New World genus Quichuana Knab, 1913 (Diptera: Syrphidae), including descriptions of 24 new species

Head: Vertical triangle black, only pollinose in front of the anterior ocellus; ocellar triangle with long, anteriorly directed, black hairs; frontal triangle black, with long, anteriorly directed, golden yellow hairs and scattered black ones medially; antenna thinly pollinose and reddish; basoflagellomere kidney shaped (Fig. 1); bf = 1.7; face extensively white pollinose, with a central shiny black stripe not reaching the antennae; face with pale hairs except for the central stripe; lower half of the occiput with white pollinosity; occiput only with golden yellow hairs.

Thorax: Scutum black with golden yellow hairs; golden yellow hairs also present on the pleuron, forming tufts on the NP and posterodorsal area of the PAPT; hairs on the PC longer than the ones on the scutum; scutellum with pale hairs; scutellum black anteriorly and dark-brown posteriorly; legs uniformly light red, except femora darker red; legs mainly with golden yellow hairs, except fine, adpressed, black hairs on the apical quarter of metafemora dorsally, and scattered black hairs on the metatibia ventrally; wings extensively microtrichose.

Abdomen: Black terga, mainly with golden yellow hairs; tergum I apparently lacking a conspicuous, moustache arrangement of hairs; hairs longer on the lateral margins than on the centre of terga; the longest hairs of the terga occurring on the anterior corner of tergum II, as in other species; sterna with pale hairs centrally.

Genitalia: Superior lobes gently serrate anteriorly, excavated apically and slightly curved backwards (Fig. 3); epandrium with surstyli expanded anteriorly at mid height (Fig. 4).

Head: Eye hairs straight and brown, slightly lighter and sparser on the lower half of eye; black vertical triangle, only pollinose on the anterior corner; ocellar triangle with long black hairs; frontal triangle black, with black hairs as long as those on the ocellar triangle and, at the eye margins, pale hairs; antenna dark; scape and pedicel with black hairs; basoflagellomere oval (Fig. 10), brownish ventrobasally; bf = 1.1; ventral part of the frontal prominence brownish; black face, with long white hairs except for a central, bare, shiny stripe extending from the bottom of the frontal prominence to the mouth edge; facial pollinosity white and restricted to a narrow line at the eye margins and a stripe extending from the eye margin to the mouth edge laterally; occiput white pollinose, with long hairs mostly white but, on the upper third of occiput, with a line of intermixed black hairs.

Thorax: Black; scutum with two medial, grey pollinose stripes extending from the anterior margin to a point halfway between the TS and the posterior margin; pale haired mainly in front of TS and black haired behind (a few fine pale hairs near wing bases); PAPT, NP, and PC with tufts of white hairs; brown scutellum, black on the lateral corners; scutellum with intermixed long pale hairs and scattered shorter black hairs; femora, tibiae, and tarsi red; metafemur darkened dorsally, but not along the full length; metatibia with a dark central ring of about a third of tibia length; pro- and mesoleg with tarsomeres 3–4 darkened dorsally and pale ventrally, metaleg with tarsomere 5 also darkened dorsally; legs with both pale and black hairs on femora, tibiae, and tarsi; femora with scattered very long hairs; cells CuP, BM, R, DM and R₄₊₅ with bare or very sparsely microtrichose areas; wings light-brown, pigmented on about the anterior third (cell R₂₊₃ very lightly pigmented).

Abdomen: Terga II–IV each with two lateral red spots covering the full length of the lateral margins of terga II–III (spots faint on tergum IV); tergum I with a moustache arrangement of white hairs with a central gap of about a third of the tergum width; terga II–IV with pale hairs except for the posterior margin that has a band of black hairs: on tergum II the length of this band is about two-thirds of the tergum length; on tergum III the band is half of the tergum length and on tergum IV the band is very narrow.

Genitalia: Superior lobes in the shape of a straight stick, nearly as long as the rest of the hypandrium (Fig. 15); epandrium with a massive surstyli, which, viewed in profile, is basally wider than high along the posterior margin (Fig. 16).

Head: Eye hairs straight, no curved apex, and light brown, slightly denser on the upper surface of eye; vertical triangle with long hairs, which are brownish-black and, on the surface immediately behind the ocellar triangle, bright yellow; frontal triangle black, with anteriorly directed hairs that are brownish black, and on the eye margins are shorter and bright yellow; basoflagellomere almost square shaped; pedicel and basoflagellomere extensively black, only narrowly reddish at the apex of pedicel and at the base of basoflagellomere; bf = 1.1; frontal prominence yellowish and thinly pollinose below antennae; face shiny black; facial pollinosity white and restricted to a narrow band on the eye margins; face with both bright-yellow and brownish-black hairs, long and down facing, except for a central bare stripe extending from the mouth edge to the bottom of the frontal prominence; occiput with bright-yellow hairs, longer on the upper and lower thirds of occiput, but also, on the upper third, with a line of intermixed, long, black hairs.

Thorax: Scutum with two medial grey pollinose stripes extending from the anterior margin, where they converge, to a point anterior to TS; long black hairs, intermixed with yellow hairs mainly on the surface anterior to TS; PP with long hairs, both black and yellow; PAPT posterodorsally, NP and PC with a tuft of black hairs; brownish-black scutellum, with black hairs as long as those on scutum, but, on the posterior margin, longer and both black (most of them) and yellow intermixed; red legs except for black coxae, trochanters, femora (narrowly red apically, even narrowerly in metafemur), and tarsomeres 3–5; protibia blackish dorsally, mesotibia extensively darkened and metatibia blackish; meso- and metatarsomeres 3–5 yellowish ventrally; posterior face of fore- and mesofemur with long erect black hairs; metafemur with long black hairs intermixed with a few pale hairs, which are both short and very long ventrally; metatibia with short black hairs; wings wholly microtrichose, very lightly pigmented; black ventral calypter, with very short black hairs centrally; calypter margins fringed with black hairs; light-red halter.

Abdomen: Tergum I black; anterior half of tergum II extensively black, but posterior half with two lateral nearly triangular red markings and a narrow red band at the posterior margin; terga III and IV red with a triangular black area anteriorly, but the anterior margin of the terga narrowly red; tergum I with a moustache arrangement of long hairs, both black and grey intermixed; anterior third of terga II–IV with yellow hairs, longer towards the lateral margins (hairs black on lateral margins of tergum II); the remainder surfaces of terga II–IV with black hairs, more backwards inclined when closer to the posterior margin; sterna I–IV with scattered long yellow hairs centrally and a few black hairs laterally, except for sternum II, with black hairs also centrally.

Genitalia: Superior lobes of hypandrium anteriorly curved, nearly rounded at apex and very finely serrated on the interior ridge of apex (Fig. 19); surstyli with a triangular process anteriorly (Fig. 18).

Head: Hairs denser on the upper surface of the eye than the lower surface; lower half of the eye with whitish hairs becoming darker towards the top of the eye; vertex and frons black; ocellar triangle and surface immediately behind with long black hairs; upper area of the frons greyish pollinose; frons with both black and light-brown hairs intermixed, although the light-brown hairs are more abundant on the upper area of the frons and the full length of the frontal eye margins; whole length of the eye margins narrowly pollinose; reddish scape and pedicel, with black hairs; basoflagellomere red, although darkened dorsally; basoflagellomere oval (Fig. 9); bf = 1.2; frontal prominence brownish below and to the sides of the antennae; frontal prominence with sparse pollinosity below the antennae; black face, with brownish white hairs and, near the mouth edge, silver white hairs; face with a central, smooth, bare area; white pollinosity restricted to a lateral arc-shaped marking, touching the eyes; genae and occiput white pollinose; occiput with white hairs and, on the upper third, with a line of long black hairs.

Thorax: Scutum ground colour black, with black pollinosity; scutum with two medial greyish pollinose stripes reaching the posterior margin of the scutum, and diverging posteriorly; scutum with brownish hairs intermixed with scattered black hairs (area above the wings and anterior part of PC with black hairs); NP, PAPT posterodorsally and PC posteriorly with conspicuous tufts of golden yellow hairs (golden yellow hairs on PC scarcer); dorsal area of AEP only with long pale hairs; anterior and posterior spiracle pale; scutellum brownish black, with long brown hairs; femora extensively black, with red on the following areas: apical quarter of the profemur, apical third of the mesofemur, and apical fourth of the metafemur; pro- and mesotibia extensively red; metatibia red at the apex and base; tarsi red; femora and tibiae mostly with pale hairs; wing lightly but conspicuously brown pigmented anteriorly, even lighter apically; wing wholly microtrichose; calypter brown centrally and black marginally.

Abdomen: Extensively dull black; tergum I with two triangular groups of golden yellow hairs obscuring the background colour of the tergum, and separated for a distance of nearly a third of the tergum width; tergum II with two very inconspicuous reddish markings laterally; tergum III with two small elongate red markings on the anterior margin; posterior margin of terga II–IV laterally fringed with conspicuous, adpressed, golden yellow hairs; terga II–IV with light-yellow erect hairs, except for a semicircular band of posteriorly directed black hairs on the posterior margin (the band of black hairs only reaches the lateral margins on tergum II); black sterna, with long pale hairs.

Head: Eye hairs pale; vertical triangle shiny black, only pollinose on the anterior corner; ocellar triangle with long black hairs; frontal triangle shiny black, with an inconspicuous dome on the upper half; frontal triangle with long, anteriorly directed, black hairs on the upper half, shorter, anteriorly directed, black hairs on the lower half, and with a line of pale hairs on the eye margins (the line of hairs does not reach the upper corner of the frontal triangle); black antenna, with thinly pollinose basoflagellomere; bf = 1.8; basoflagellomere elongate; black face, with a central shiny stripe not reaching the antennae and two wide lateral shiny stripes, with the rest of the face pollinose.

Thorax: Black scutum with posteriorly directed pale hairs; pleuron pale haired, without conspicuous tufts of hairs; anterior and posterior spiracles pale in colour; scutellum with pale hairs, longer on the posterior margin; legs black, except for the red apex of the femora (red area larger on metafemur), full tibiae, and tarsi; legs with pale hairs except for the apicodorsal area of the femora bearing short, black, adpressed hairs (the black hairs cover a larger area on the metafemur), and the apical third of the metafemur ventrally bearing spiny black hairs; wings extensively microtrichose, except for the cells BM and CuP, with large bare areas; wings lightly brown pigmented anteriorly.

Abdomen: Black, extensively pale haired, with hairs on the lateral margins of terga II–IV longer than those positioned centrally on these terga (hairs on the lateral margins straight and with the maximum length on tergum I posteriorly and tergum II anteriorly); tergum I with a moustache arrangement of silver white hairs; terga II–III with a very narrow, white pollinose band on the posterior margin; sterna with long white hairs.

Head: Eye hairs straight and brown, slightly lighter on the lower half of the eye and denser towards the upper half; vertical triangle black with white pollinosity on the anterior corner and, on the ocellar triangle, long black hairs; frontal triangle black with long black hairs and, on the eye margins, a line of pale hairs; antenna pale; basoflagellomere oval and darker than scape and pedicel; bf = 1.2; scape and pedicel with black hairs; ventral area of the frontal prominence red, with sparse pollinosity; black face, with long white hairs except for a bare, central, shiny stripe from the mouth edge to the bottom of the frontal prominence; facial pollinosity white, and restricted to a narrow line at the eye margin and a stripe extending from the eye margin to the mouth edge; occiput white pollinose, with long white hairs and, on the upper third, a line of long black hairs.

Thorax: Black; scutum with two medial, approximated, brownish pollinose stripes extending on the anterior four-fifths of the scutum; scutum with both yellow and black hairs intermixed; black hairs denser on the half of the scutum behind TS and also occurring on the anterior area of the PC and above the wings; PAPT posterodorsally, NP and PC with tufts of golden yellow hairs; brown scutellum, blackish on the lateral corners; scutellum with yellow hairs and, centrally, scattered black hairs intermixed; basal half of the pro- and mesofemur black; basal three-quarters of the metafemur black; pro- and mesotibia red; metatibia black on about the central third; all tarsi red except for the metatarsomeres 4–5 dorsally; femora and tibiae with both pale and black hairs; protarsomeres 3–5, mesotarsomeres 2–5, and metatarsus with black hairs dorsally; anterior third of the wing faintly brown pigmented; wing extensively microtrichose, except for some areas with lower density of microtrichia on cells R and CuA1.

Abdomen: Terga II–III each with two lateral red spots, which cover less than half of the surface of each tergum, including the full-length of the lateral margins of tergum II; tergum IV with two very faint, small, pale spots; terga II–IV black pollinose, except for a narrow, shiny, posterior band, slightly larger in tergum IV; tergum I with a moustache arrangement of golden yellow hairs, with a central gap of about a third of the tergum width; terga II–IV yellow haired, except for the posterior margin, which has a band of black hairs on the posterior half of the tergum or less; posterior margin of terga II–III laterally fringed with adpressed yellow hairs (Fig. 77).

Genitalia: Hypandrium with sickle-shaped superior lobes (Fig. 25); surstyli tapering towards the apex, round apically (Fig. 26), and with a round expansion on the inner side (Fig. 27).

Head: Eye hairs brownish, denser on the upper surface; black vertical triangle, only pollinose on the anterior corner; vertical triangle with long black hairs; frontal triangle black, with long, anteriorly directed, black hairs and, at the eye margins, a line of yellow hairs; antenna red; basoflagellomere slightly darkened; bf = 1 (Fig. 14); ventral part of the frontal prominence red; black face with long yellow hairs except for a central bare stripe; facial pollinosity white and restricted to a narrow line at the eye margins and a very faint stripe extending from the eye margin to the mouth edge laterally; occiput white pollinose, with white hairs, but with a line of intermixed long black hairs on the upper third of occiput.

Thorax: Black scutum with two medial, approximated, parallel, brownish pollinose stripes, extending from the anterior margin of scutum to a point behind TS; scutum with intermixed black and yellow hairs, although black hairs more abundant on the scutum surface behind TS; pleura pollinose; PAPT, NP, and PC with a dense group of long, mainly black hairs, although anterior part of PC with yellow hairs; red scutellum, black on lateral corners; scutellum with both black and yellow hairs, longer on the posterior margin; pro- and mesotibia red; metatibia with some areas conspicuously darkened; tarsomeres 3–5 black dorsally; femora extensively black, only narrowly red at the apex; profemur mainly with black hairs, only a few very short pale hairs at the apex; mid- and metafemur, all tibia, and tarsus with both black and yellow hairs; wings faintly brown pigmented on the anterior half; wings extensively microtrichose; calypter black.

Abdomen: Tergum I with a moustache arrangement of yellow hairs, which does not obscure the background colour of the tergum; terga II–IV red, or nearly so; terga II–III with a black area anteriorly; at least posterior third of terga II–IV with a band of black hairs extending forwards along the lateral margins, but not reaching the anterior margin; terga III–IV with scattered long hairs on lateral margins.

Genitalia: Superior lobes anteriorly curved in profile view (Figs 32, 36); interior area delimited by the curved lobes in the shape of a horseshoe, viewed in profile (Figs 32, 36); base of the surstyli with a triangular process anteriorly and, posteriorly, with long hairs (Fig. 37).

Head: Eye hairs whitish and sparse on the lower half of the eye, and brown and denser towards the upper half; vertical triangle black with white pollinosity on the anterior corner and, on the ocellar triangle, long black hairs; frontal triangle black, with long black hairs except for the line of brownish white hairs on the eye margins; dorsal surface of the frontal triangle brownish near the antennae; antenna with arista red; scape black; pedicel and basoflagellomere blackish red; basoflagellomere oval; bf = 1.1; scape and pedicel with black hairs; ventral surface of the frontal prominence reddish, with white pollinosity below the antennae; black face, with most of the hairs brownish white; face with a bare, central, shiny stripe from the mouth edge to the bottom of the frontal prominence; facial pollinosity white and restricted to a narrow line at the eye margin and a stripe extending from the eye margin to the mouth edge laterally; occiput white pollinose, with white hairs and, on the upper third, scattered black hairs intermixed.

Thorax: Black; scutum with two medial, approximated, greyish pollinose, tapering stripes, faintly reaching the posterior margin of the scutum (the rest of the scutum covered with darker pollinosity); scutum with reddish hairs and, above the wings and the anterior area of the PC, black hairs; NP, PAPT posterodorsally, and PC posteriorly with conspicuous tufts of golden yellow hairs; scutellum brown, nearly black on the lateral corners; scutellum with long red hairs and a few long black hairs intermixed; anterior and posterior spiracles pale; apical half of pro- and mesofemur with red markings; metafemur black, with red markings on the apical third; pro- and mesotibia extensively red; protibia with a blackish ring near the apex; metatibia red on the basal half; tarsi red, except for the darker metatarsomeres 3–5; femora and tibiae with both pale and dark hairs; wing entirely microtrichose, very lightly brown pigmented anteriorly.

Abdomen: Terga II–IV each with red markings laterally and posteriorly; posterior half of tergum II with a band of black hairs reaching the lateral margins; posterior half of tergum III with black hairs medially; tergum I with a moustache arrangement of golden yellow hairs nearly obscuring the background colour of the tergum, with a central gap of a third of the tergum width; sterna black with long pale hairs, except for scattered black hairs on sternum IV.

Genitalia: Hypandrium similar to that in Q. bromeliarum sp. nov. (Fig. 25); base of the surstyli about twice wider than the apex; surstyli gently concave posteriorly and, from mid height, abruptly tapering anteriorly (Fig. 38); inner side of the surstyli with a triangular expansion (Fig. 39).

Head: Frons brown pollinose, with both black and light-brown laterally directed hairs intermixed; frons with a line of shorter, light-brown hairs on the eye margins; hairs on the dorsal area of the frontal prominence anteriorly directed; antenna red, only blackish on the basoflagellomere dorsally; basoflagellomere oval (Fig. 11); bf = 1.3; frontal prominence ventrally black, except for the red area surrounding the antennae; black face with white hairs, except for a central shiny bare stripe; the rest of the face covered with very sparse white pollinosity, except for the dense pollinosity on the eye margins; gena with a lateral densely pollinose stripe from the eye margins to the mouth edge.

Thorax: Scutum with two brown pollinose stripes faintly reaching the posterior margin of the scutum; scutum extensively pale haired, but with a few short black hairs on the area above the wing insertion; PAPT posterodorsally, NP, and PC with dense and conspicuous tufts of whitish yellow hairs; AEP with whitish yellow hairs; brown scutellum, black on the lateral corners; scutellum covered only with long yellowish hairs; anterior and posterior spiracles pale; the apical two-thirds of pro- and mesofemur, the full length of the pro- and mesotibia, and the pro- and mesotarsus red; apical third of metafemur red; metatibia red with black markings on the apical two-thirds; metatarsus red; legs extensively pale haired, except for black hairs present on the following areas – the apical half of the profemur posteriorly, the protibia (scattered black hairs), the apex of the mesofemur posteriorly, the apical third of the metafemur ventrally, the apex of the metafemur dorsally, most of the length of the metatibia ventrally, and the apical half of the metatibia dorsally; wings extensively microtrichose; wings very lightly brown pigmented anteriorly.

Abdomen (73–79): Tergum I with a moustache arrangement of whitish yellow hairs, with a central gap; terga II–IV each black with two lateral red markings, which extend from the anterior to the posterior margins in terga II–III, and only reach the anterior margin in tergum IV; terga II–III narrowly red on the full length of the posterior margin, even narrower in tergum III; terga II–IV extensively brownish pollinose except for the shiny band on the posterior margin of tergum IV; tergum V shiny except for an anterior narrow dull band; all other terga extensively covered with yellowish white hairs, except for black hairs on the following areas – posterior black areas of terga II–IV centrally and the posterior half of the lateral margins of tergum II (black hairs also on the posterior corners of tergum II); posterior margins of terga II–IV laterally fringed with adpressed yellowish white hairs; tergum V with long black hairs, but pale hairs anteriorly; sterna with long pale hairs; sternum V with scattered black hairs intermixed with pale hairs.

Head: Brown pollinose frons, with black hairs and, at the eye margins, pale hairs; basoflagellomere oval; bf = 1.2; frontal prominence reddish below and to the sides of the antennae; lower part of frontal prominence thinly white pollinose; white-haired face, with two lateral white pollinose stripes extending from the mouth edge to the frontal prominence (the pollinose stripes extend along the eye margins and taper at the upper end, which connects with the sparse pollinosity of the lower part of the frontal prominence); face with a central, bare, shiny stripe.

Thorax: Scutum with two medial, wide, brown pollinose stripes faintly reaching the posterior margin of the scutum; PAPT posterodorsally, NP, and PC with conspicuous tufts of white hairs; brown scutellum, black on lateral corners; scutellum with long pale hairs only; pro- and mesotibia, tarsi, and apex of femora red; metaleg blackish, except for the extreme apices of the femora and tibiae, the tarsus ventrally, and tarsomeres 1–2 dorsally, which are red; legs with pale hairs except for the following areas, which have black hairs – apical half of profemur posteriorly, apical third of metafemur posteriorly and ventrally, and at the apex of metafemur dorsally; wings extensively microtrichose except for some small bare areas in cells BM and CuP, and at base of the anal lobe; wings very lightly brown pigmented anteriorly.

Abdomen: Terga II–III each with two lateral red markings, each connecting the anterior and posterior margin in tergum II; tergum III with two additional, narrow, red bands on the posterior margin; tergum IV black; terga II–V extensively coated in brownish pollinosity, except for a posterior band, which is shiny; tergum I with a moustache arrangement of white hairs with a central gap; terga II–IV mainly with white hairs, but with scattered black hairs posteriorly on the following areas – black parts of terga II–III centrally and lateral margins of the posterior half of tergum II; posterior margin of terga II–IV laterally fringed with conspicuous, adpressed, white hairs (Fig. 77); tergum V with long black hairs posteriorly; sterna with long pale hairs centrally.

Head: Eye hairs slightly lighter on the lower half of the eye; ocellar triangle with black hairs; frons with sparse brownish pollinosity; frons with black and golden yellow hairs intermixed dorsally, and black hairs ventrally and on the dorsal area of the frontal prominence; eye margins with golden yellow hairs; antenna dark red, especially darkened on the scape and the base of the pedicel; ventral area of the frontal prominence brownish, with whitish pollinosity; face black, with long golden yellow hairs, except for a central, bare, shiny stripe from the bottom of the frontal prominence to the mouth edge; gena with a stripe of white pollinosity from the mouth edge to the eye margin; occiput wholly pollinose and with long golden yellow hairs, and, on the upper third, a line of long black hairs.

Thorax: Black; scutum with two medial brownish pollinose stripes reaching the posterior margin of the scutum; scutum yellow haired, except black haired behind the TS and on the area above the wings; NP, PAPT, and PC with tufts of golden yellow hairs; scutellum dark brown, black on lateral corners; anterior half of the scutellum black haired centrally and posterior half yellow haired; all femora extensively black; apical half of the pro- and mesofemur red anteriorly; apical half of the protibia with blackish markings, the remaining area red; mesotibia red; apical two-thirds of the metatibia blackish, the remaining area red; tarsi red, except for the darkened dorsal area of metatarsomeres 4–5; femora, tibiae, and tarsi mostly with golden yellow hairs, but at least with scattered black hairs or setae; wings wholly microtrichose and brown pigmented on the anterior half.

Abdomen: Black, without conspicuous pale markings; most of the surface of terga II–V covered with sparse brownish pollinosity; tergum I with a moustache arrangement of golden yellow hairs, with a central gap of about a third of the tergum length; posterior margin of terga II–IV laterally fringed with adpressed golden yellow hairs; posterior half of terga II–IV with a medial nearly circular patch of black hairs; lateral margins of the posterior half of tergum II also with scattered black hairs; tergum V mainly black haired.

Head: Eye hairs straight, no curved apex, brown and dense on the upper half of the eye, and slightly lighter and sparser on the lower half; vertical triangle black with white pollinosity on the anterior corner; ocellar triangle with long black hairs; frontal triangle black, with long black hairs centrally and pale hairs on the eye margins; antenna extensively pale; bf = 1; ventral area of the frontal prominence brownish with sparse pollinosity, and with a tapering pollinose stripe from the eye margin towards the antennae; black face with long white hairs, except for a bare central shiny stripe from the frontal prominence to the mouth edge; gena with a stripe of white pollinosity extending from the eye margin to the mouth edge; occiput pollinose, with long white hairs and a line of long black hairs intermixed on the upper third.

Thorax: Black; scutum with two grey pollinose stripes extending inconspicuously beyond the TS; scutum with yellow hairs and, on the half of the scutum behind the TS, black hairs intermixed; area above the wings bearing black hairs; PAPT posterodorsally and NP with tufts of golden yellow hairs; PC mainly with long black hairs; scutellum dark brown, black on lateral corners; scutellum with yellow hairs and scattered black hairs intermixed; all femora extensively black, paler on the apical quarter; pro- and mesotibia red; metatibia red, with a central dark ring and on the apical two-thirds dorsally; all tarsi red, slightly darkened on tarsomeres 4–5 dorsally; all femora and tibiae with both yellow and black hairs; tarsi mainly yellow haired but with at least a few black hairs or setae on tarsomere 5; wing extensively hyaline, only dark-brown pigmented on some areas of the very basal cells; wing microtrichose, except for some bare areas on cells R, BM, and CuP.

Abdomen: Terga II–IV each with two red markings extending on the full lateral margins and covering more than half of the tergum surface; tergum I with a moustache arrangement of golden yellow hairs, with a gap of about a third of the tergum width; terga II–IV with yellow hairs anteriorly, but black hairs on the posterior half or less.

Genitalia: Hypandrium with comma-shaped superior lobes (Fig. 40); epandrium with surstyli wide at base, tapering towards the apex, slightly curved backwards in profile (Fig. 41); surstyli with an excavation and a triangular process on the inner side (Fig. 42).

Head: Frontal triangle with a conspicuous dome on the upper half (see head in lateral view), and with long black hairs centrally and yellow hairs on the eye margins (Fig. 81); dorsal surface of the frontal prominence swollen; scape conspicuously shorter than pedicel or basoflagellomere (Hull, 1946: 16, fig. 18); bf = 3; black face, with narrow lines of pollinosity from the antennae to the mouth edge, and delimiting a central, shiny, black stripe and two lateral stripes (Hull, 1946: 16, fig. 11; 17, fig. 24).

Thorax: All hairs on PP yellow; scutum with two medial, greyish pollinose stripes extending to the anterior three-quarters of the scutum; scutum with short, posteriorly directed, yellow hairs and a few much longer black hairs intermixed on the half of the scutum behind the TS; PC and area just above the wings with black hairs, shorter than the black hairs of the rest of the scutum; pleuron with yellowish white hairs, tufted on NP, PAPT, and PC; metaepisternum with a few short, pale hairs; metafemur conspicuously swollen, reddish on the apical third; pro- and mesofemur black to dark brown; tibiae and tarsi red to reddish brown; halteres dark brown.

Abdomen (80–83): Black; tapering from tergum I to about the middle of tergum II, and then widening progressively until tergum IV; terga II–III with a conspicuous yellowish white pollinose band on the posterior margin; tergum I with a moustache arrangement of yellowish white hairs; terga I–II with long yellowish white hairs on the lateral margins (with the longest hairs on tergum II); terga II–IV with yellowish white hairs and scattered black hairs intermixed; hairs on the posterior margin of terga II–IV longer than those on the rest of the tergum, yellow on terga II–III, and black on tergum IV.

Head: Eye hairs brownish; vertical triangle pollinose on the anterior corner; ocellar triangle with long, anteriorly directed, black hairs; frontal triangle and face extensively shiny black, only narrowly white pollinose on the eye margins; antenna thinly pollinose, black, narrowly red at the very base of the basoflagellomere; basoflagellomere nearly round (Fig. 43); bf = 1.25; face pale haired, except for a central bare stripe extending from the antennae to the mouth edge; occiput with yellow hairs and, on the upper half, long black hairs intermixed; lower half of the occiput pollinose.

Thorax: Black; PP with both black and pale hairs and a faint pollinose marking; scutum thinly grey pollinose, with two faint, medial, white pollinose stripes extending beyond the TS; scutum extensively pale haired, but with black hairs on the area above the wing insertion and between the wings; PC black haired; AA with a few fine yellow hairs ventrally; PAPT posterodorsally with a tuft of long yellow hairs; AEP with black hairs; katepisternum with fine yellow hairs; scutellum with long pale hairs; legs black and extensively black haired, but with pale hairs at least on the metafemur; metafemur slender; calypteres reddish; wings extensively microtrichose and very faintly pigmented in the basal part of the anterior margin.

Abdomen: Black; terga I–IV with pale hairs and, on terga II–IV posteriorly, black hairs; tergum I with a moustache arrangement of sparse light-yellow hairs; lateral margins of terga II–IV with long light-yellow hairs; sterna with long light-yellow hairs centrally.

Genitalia: Superior lobes short and nearly square in lateral view (Fig. 45); epandrium with surstyli round at the apex, slightly expanded anteriorly (Fig. 46).

Head: Eyes with white hairs, except for the brownish hairs on upper third of the eye; ocellar triangle with black hairs; frontal triangle with long, black hairs centrally and pale hairs laterally; antenna black, except for red arista; basoflagellomere nearly rectangular, round at the apex (Fig. 2); bf = 1.9; face pollinose, with a central, shiny, black stripe, not reaching the antennae, and two lateral stripes slightly narrower than the central stripe (lateral stripes with sparse pollinosity not obscuring the background colour); frontal prominence pollinose ventrally; face with silver, white hairs.

Thorax: Scutum with two pollinose stripes tapering towards the apex, each stripe as wide as half the width of the basoflagellomere; pollinose stripes extending on the anterior two-thirds of the scutum; scutum with long, yellowish white, posteriorly directed hairs, and a few short black hairs above the wings; pleural hairs pale, with some reduced tufts of hairs on PAPT posterodorsally and NP; PC with long, yellowish hairs; scutellum dark brown, with long pale hairs; pro- and mesofemur, tibia, and tarsus dark brown, although tibia and tarsus lighter; metafemur black, with reddish areas apically; metatibia extensively dark brown, red at the very apex; femora and tibiae mainly pale haired; dorsal surface of the apical third of metafemur with short, black, adpressed hairs and spiny black hairs on the ventral surface of the apical third; tarsi with short, black, adpressed hairs dorsally; wings weakly brown pigmented on the surface, anterior to vein R₄₊₅ (cell C even with lighter pigmentation than that on the rest of the pigmented surface); wing with areas bare of microtrichia on cell R posterior to spurious vein, on most of the basal third of C, most of the basal two-thirds of BM, and most of the basal third of CuP; alula with a semicircular area bare of microtrichia anteriorly; reddish white calypteres with blackish margins; halteres reddish.

Abdomen: Slender and tapering towards the apex; terga black; lateral surface of tergum I with laterally directed hairs; terga with whitish yellow hairs, except for the black hairs on the posterior third of tergum II, posterior two-thirds of tergum III, and posterior third of tergum IV; sterna dark brown.

Genitalia: Similar to that of Q. angustiventris but with superior lobes shorter and more developed on the anterior surface (Fig. 8).

Head: Upper quarter of eyes with dense, brownish white hairs; occasional white hairs at the middle of the eyes; lower margin of eyes with sparse, straight, white hairs; vertical triangle with long, black hairs; a row of long, golden yellow hairs just posterior to the ocellar triangle; frontal triangle black, with long, anteriorly directed, golden yellow hairs laterally and scattered, long, black hairs centrally; antenna elongate, basoflagellomere 1.3 × longer than pedicel and pedicel 2.7 × longer than scape; ventral surface of frontal prominence yellowish laterally; face black, with lateral, white pollinosity delimiting a central, rhomboidal, shiny area; long, bright-yellowish white hairs on both the pollinose and shiny surfaces of face, except for a median, narrow, bare stripe; occiput with long, golden yellow hairs, shorter and paler when further down the occiput (upper section of occiput with a row of black setae intermixed with yellow hairs); from the midpoint of the occiput to the lower end, hairs progressively longer and sparser.

Thorax: Scutum with two grey pollinose stripes, nearly reaching the posterior margin of scutum; scutum with golden yellow hairs as long as those on scutellum; NP, PAPT posterodorsally, and PC with a tuft of golden yellow hairs; scutellum brownish black, with long, golden yellow hairs; proleg red, except for the black coxa and trochanter; profemur narrowly yellow apically; meso- and metaleg red, except for the black coxa, trochanter, and basal two-thirds of the femur; pro- and mesofemur and tibia with long, yellow hairs posteriorly; tibiae with black hairs dorso-apically (in mesoleg also ventro-apically); tarsi with short black hairs dorsally; metatrochanter with a conspicuous process bearing apically a row of black setae (Fig. 50); metatibia inflated towards the apex, constricted ventrally and with an apical, short process bearing black, adpressed hairs (Fig. 49); metafemur with long yellow hairs, longer on anterior surface; a few short, spiny, black setae on ventral third of femora; metatarsus yellow, with black setae dorsally; wings extensively microtrichose, with small, bare areas in cells bc, C, and CuP; wings with light-brown pigmentation in cells SC and R₁, and in cells R₂₊₃ and R anteriorly; ventral calypteres white centrally, but black with short, black hairs in the margin; halteres whitish yellow apically, brown on the basal third.

Abdomen: All terga dull, bronzy black, except for shiny surface of terga I and II anteriorly; tergum I with a moustache arrangement of long, golden yellow hairs; tergum II only with yellow hairs, very long on the anterior section of lateral margins; tergum III only with golden yellow hairs; tergum IV with golden yellow hairs and scattered black hairs posteriorly; tergum V with both whitish yellow hairs and scattered black hairs; sternum I with long white hairs; sterna II and III with long yellowish white hairs; sternum IV with very few lateral hairs, both pale and black intermixed.

Genitalia: Superior lobes short and more or less conical; secondary processes of hypandrium developed with respect to those in the genitalia of species such as Q. angustiventris; surstyli slender, narrowed at the base, but widening towards the apex; base of the surstyli with a tooth in the interior side.

Head: Eye hairs darker and denser on the upper half; vertical triangle shiny black, with long black hairs on the ocellar triangle; frontal triangle extensively shiny black, with pollinosity on the upper corner and on the eye margins; frontal triangle with long, anteriorly directed, black hairs; antenna thinly pollinose, blackish, with arista reddish; basoflagellomere as long as the pedicel or slightly shorter (Fig. 53); bf = 1.5; face extensively shiny black, only narrowly white pollinose on the eye margins; face with long white hairs except for a central bare stripe from the antennae to the mouth edge.

Thorax: Scutum and pleuron extensively black haired; scutum with two medial white pollinose stripes extending to the TS or slightly beyond; PC with long black hairs; AEP with long black hairs at least dorsally; anterior spiracle blackish and posterior spiracle whitish; dark-brown scutellum with long black hairs centrally and longer light-brown hairs posteriorly; legs extensively black, only tibiae and apical tarsomeres dorsally paler; legs with black hairs, except for scattered pale hairs on tarsi; wings anteriorly pigmented and bare of microtrichia on extensive areas of cells BM and CuP, and smaller areas of cells DM and CuA1; calypteres black.

Abdomen: Abdomen shiny black except for an area on the anterior margin of tergum II; tergum I with a moustache arrangement of laterally directed whitish yellow hairs; terga II–IV extensively black haired; tergum II with long black hairs on the lateral margins, decreasing in length backwards; sterna with long white hairs centrally.

Genitalia: Same as that in Q. cincta (Figs 28, 29).

Head: Frontal triangle with long anteriorly directed black hairs and, on the eye margins, a line of light-yellow hairs; scape, pedicel, and basoflagellomere red, darkened dorsally; arista red; bf = 1.3; ventral area of the frontal prominence extensively red and thinly white pollinose on the area surrounding the antennae; face shiny black, with light-yellow hairs, sculptured laterally, but bare and smooth centrally, from the frontal prominence to the mouth edge; face with white pollinosity on the eye margins; gena with a stripe of white pollinosity extending from the eye margin to the mouth edge.

Thorax: Scutum with two medial brown pollinose stripes extending on the anterior four-fifths of the scutum (posterior section of each stripe tapering towards the apex); scutum extensively covered with golden yellow hairs, but with scattered black hairs intermixed on the area above the wing insertion; PAPT posterodorsally, NP, and PC with conspicuous tufts of golden yellow hairs; AEP with golden yellow hairs; anterior spiracle reddish white; posterior spiracle white; scutellum brown, black on the lateral corners; scutellum only with long golden yellow hairs; tibia, tarsi, and apex of the femora red (metalegs darker, with nearly black markings on tibia); legs extensively pale haired, except for black hairs on the apical half of the profemur posteriorly, the apical third of the mesofemur posteriorly, and the apical third of the metafemur dorsally and ventrally; wings extensively microtrichose, very weakly brown pigmented on the basal part of the anterior margin.

Abdomen: Terga II–IV each with two lateral red markings, extending from the anterior to the posterior margin (narrowly on tergum IV posteriorly), and on the full length of the lateral margins; terga II–III red on the full length of the posterior margin; tergum I with a moustache arrangement of golden yellow hairs; terga extensively covered with golden yellow hairs; terga with black hairs on the posterior margin centrally and on the lateral margins of the posterior half of tergum II; posterior margin of terga II–IV laterally fringed with golden yellow hairs; sterna with long golden yellow hairs centrally and, on sternum IV posteriorly, black hairs.

Genitalia: Superior lobes as illustrated in Figure 56; surstyli broadly triangular, excavated basally, round at the apex (Fig. 57).

Head: Eye hairs light brown and denser on the upper third, otherwise white; ocellar triangle with black hairs; frons with black hairs and, dorsally, intermixed with pale hairs; hairs on frons dorsally laterally directed; dorsal area of the frontal prominence only with black anteriorly directed hairs; frons with golden yellow hairs only on the eye margins, and not extending beyond the ocellar triangle; frons with sparse brownish pollinosity medially and, on the pale-haired areas, laterally; scape dark brown; pedicel light brown; basoflagellomere oval, nearly black; bf = 1.3; ventral area of the frontal prominence brownish and white pollinose below the antennae; face black with long golden yellow hairs, except for a bare central shiny stripe extending from the frontal prominence to the mouth edge; gena with a white pollinose stripe from the eye margin to the mouth edge; occiput with white pollinosity on the eye margins; occiput with long golden yellow hairs and, on the upper third, a line of long black hairs.

Thorax: Black; scutum with two brownish pollinose approximated stripes, faintly reaching the posterior margin, yellow haired except for a few black hairs behind the TS and on the area above the wings; PAPT posterodorsally, NP, and PC with tufts of golden yellow hairs; scutellum dark brown, black on the lateral corners; scutellum mainly pale haired, but with black hairs centrally; profemur red except for the black basal two-thirds posteroventrally; mesofemur red; metafemur black except for the red apical third; pro- and mesotibia red; metatibia red but with darkened markings on the apical half; tarsi red; femora and tibiae mostly yellow haired, but always with at least a few black hairs or setae; wings extensively microtrichose, only sparsely microtrichose or bare in narrow areas close to the veins; anterior half of the wing with light-brown pigmentation.

Abdomen: Terga II–III each with two lateral red markings; tergum II with the red markings triangular-shaped and extending on the full length of the lateral margins; more than half of the surface of tergum II black; red markings on tergum III touching all the margins of the tergum; tergum I with a moustache arrangement of golden yellow hairs, with a gap of about a third of the tergum width; terga II–IV with golden yellow hairs, but black hairs posteriorly and laterally; posterior margin of terga II–IV posteriorly fringed with adpressed yellow hairs; tergum V mostly black haired, but with yellow hairs posteriorly.

Head: Ocellar triangle with black hairs; black frons, with whitish hairs and, posteriorly, a few black hairs; frons with white pollinosity medially; antenna black except for the red arista; basoflagellomere elongate (Fig. 54); bf = 2; face with a central shiny stripe and two lateral shiny stripes outlined with white pollinosity; gena with a thinly pollinose stripe, extending from the eye margin to the mouth edge; occiput white pollinose next to the eye margins, white haired and with a line of long black hairs on the upper third.

Thorax: Black scutum, covered with sparse greyish pollinosity and two medial approximated stripes of denser pollinosity extending as far as three-quarters of the scutum; scutum white haired in front of the TS and mainly with short black hairs behind; area above the wings and anterior area of the PC black haired; pleuron black; PAPT posterodorsally, NP, and PC with sparse groups (non-tufted) of long pale hairs; scutellum shiny, copper, white haired, and with a few black hairs; femora black, except for a short reddish section at the apex; tibiae black with brownish markings and a short reddish section at the base; tarsi black dorsally, except for the brownish basotarsomere; all legs mainly pale haired, with a few black hairs on some areas of femora, tibiae, and tarsi; wings extensively microtrichose; anterior third of the wing lightly brown pigmented; calypteres white except for the black margins and the marginal hairs.

Abdomen: Black; tergum I with a moustache arrangement of white sparse hairs; tergum II extensively covered with white hairs and scattered black hairs at the posterior margin; tergum III extensively black haired, only with white hairs on a narrow band of the anterior margin; terga IV–V extensively white haired, but with scattered black hairs intermixed.

Head: Eye hairs lighter and shorter further down the eye; ocellar triangle with long, black hairs; vertical triangle with pollinosity restricted to the anterior corner; frontal triangle shiny black, with long, black hairs centrally, and whitish hairs on the eye margin; basoflagellomere elongate (bf = 2.5–3, N = 8), nearly 2 × longer than the pedicel; basoflagellomere and pedicel pollinose; basoflagellomere black to dark brown; face pollinose, with a central, shiny stripe and two lateral, shiny stripes wider than the central stripe; lower part of the face with long, pale hairs in a beard arrangement, conspicuously exceeding the mouth edge (Figs 75, 76); facial hairs, including the ‘beard’ hairs, silver white to yellowish white.

Thorax: Only with yellow to whitish hairs; scutum with two pollinose, narrow, approximated stripes, diverging slightly posteriorly; NP, PAPT posterodorsally, and PC with tufts of hairs; legs extensively yellow, except for the dark coxae, trochanters, and restricted parts of the metafemur, tibiae, and tarsi; wings extensively microtrichose; alula microtrichose, except for a bare semicircular area on the anterior margin.

Abdomen: Tergum I with a moustache arrangement of usually yellow hairs; terga II–IV yellow haired, except for the posterior margin that has a semicircular band of black hairs; on terga II–III the length of this band is a quarter of the tergum length, and on tergum IV it is about a half of the tergum length; lateral margins of terga II–IV only with pale hairs.

Genitalia: Virtually the same as that in Q. angustiventris (Fig. 5–7).

Holotype: Basoflagellomere elongate (bf = 2.2); face with two lateral, wide, shiny stripes; face with silver white hairs, absent on the central, shiny stripe; vertex with black hairs around the ocellar triangle; wing very weakly pigmented, almost hyaline; laterally directed, densely-grouped, yellow hairs are visible on the left part of tergum I. Hull (1946) provides some notes based on the damaged type, including a mention of the abdominal pilosity, but there is no reference to tergum II. Hull (1946) states that the black hairs are restricted to the posterior quarter of terga III and IV, and tergum V has yellow hairs.

Additional features based on other females examined: Frons with an anteriorly directed chevron of pollinosity; chevron with erect, usually silver white to yellow hairs; area between the vertex and the V-shaped band with erect, dark-brown to black hairs; dorsal side of the frontal prominence with anteriorly directed, pale hairs, sometimes intermixed with dark-brown to black hairs; basoflagellomere elongate (bf = 2.1–2.4, N = 7); hairs on face of the same colour as those in the holotype; thorax extensively white to yellow haired, except for the short, black hairs above the wings and, occasionally, the black hairs on PC anteriorly and on the area immediately posterior to TS; NP, PAPT posterodorsally, and PC with not very dense tufts of hair; microtrichia in alula as that in males; tergum I pale haired; posterior margin of terga II–IV with black hairs extending on the same, or about the same, proportion as those in males.

Head: Ocellar triangle with pale hairs; frons with an anteriorly directed chevron of pollinosity; frons with sparse, anteriorly and laterally directed, golden yellow hairs; dorsal surface of frontal prominence with hairs conspicuously longer than those on frons; antenna black; basoflagellomere straight ventrally and curved dorsally (Fig. 20); bf = 1.5; face white pollinose, except for a very narrow central, shiny stripe not reaching the antennae nor the mouth edge; face with golden yellow hairs, bare on the central stripe; occiput with white pollinosity on the lower two-thirds.

Thorax: Scutum black, covered mainly in short, pale hairs; scutum with two conspicuous, white pollinose stripes extending beyond the TS; NP and PAPT posterodorsally with tufts of white hairs; hairs on PC longer than those on the scutum; scutellum dark brown, with short, pale hairs; legs black, except for tibiae, tarsi, and apical third of femora, which are darker, especially in metalegs; apicoventral half of metafemur with black hairs; ventral surface of mesotibia with scattered black hairs; wings brown, pigmented anteriorly and extensively microtrichose, except for some bare areas on cells BM, CuP, and alula; calypteres white, but with dark-brown margin.

Abdomen: Tapering towards the apex; terga black; terga I–IV mostly pale haired; tergum I without a conspicuous moustache arrangement of hairs; anterior section of the lateral margins of tergum II with long, white hairs, some of them wavy at the tip (the longest hairs at each of terga III–IV also occur on the anterior section of the lateral margins of terga); tergum V with scattered black hairs posteriorly; sterna with white hairs, shorter on sterna III–IV.

Head: Eye hairs brown, becoming lighter and sparser ventrally; black vertical triangle with white pollinosity only on the anterior corner; ocellar triangle with long black hairs; frontal triangle black with long black hairs and, on the eye margins, a line of pale hairs; antenna blackish red, nearly black at the apex of pedicel and at the base of basoflagellomere; basoflagellomere oval; bf = 1.2; arista red, darkened towards the apex; frontal prominence red and pollinose below antennae; frontal prominence with a faint, pollinose, tapering stripe extending from the eye margins to the antennae; black face with long white hairs except for a central bare stripe from the mouth edge to the bottom of the frontal prominence; face with white pollinosity on the eye margins and a stripe of white pollinosity from the eye margin to the mouth edge laterally; occiput white pollinose, with long white hairs, but also, on the upper third, with a line of intermixed black hairs.

Thorax: Black; greyish pollinose scutum with two medial, approximated, brownish pollinose stripes, stripes extending to about three-quarters the length of the scutum; scutum, including area immediately above wings, with intermixed red and black hairs, although black hairs more abundant on the surface posterior to TS; PAPT posterodorsally, NP, and PC with non-tufted groups of long hairs, intermixed red and black; brown scutellum, blackish on lateral corners; scutellum with intermixed red and black hairs; all femora extensively black, only lighter or yellow apically; basal half of pro- and mesotibia yellow, but apical half with some areas yellow; metatibia black, with some yellow areas at apex and base; pro- and metatarsomeres 2–5 and mesotarsomeres 3–5 black, lighter ventrally; femora, tibiae, and tarsi with both black and yellow hairs intermixed; wings extensively microtrichose, with some areas bare or with sparser microtrichia in cells BM, R, and CuA1; wings very lightly brown pigmented on the anterior half; veins on the anterior third of wing light brown.

Abdomen: Black; terga II–IV extensively brownish pollinose, with a short shiny band posteriorly (pollinosity of the same colour as that of the two medial stripes on scutum); tergum I with a moustache arrangement of reddish hairs not obscuring the background colour of tergum, and with a central gap of about half of the tergum width; terga II–IV mostly with red hairs, but also with black hairs roughly on the shiny parts of all terga, and along most of the length of the lateral margins on tergum II.

Genitalia: Superior lobes of hypandrium wedge shaped, viewed in profile (Fig. 17); surstyli with a triangular process anteriorly (Fig. 18).

Head: Eye hairs straight, not curved apically, and brown, but lighter and sparser lower down; vertical triangle black, only pollinose in front of the anterior ocellus; ocellar triangle with long black hairs; frontal triangle black, with long black hairs and, at the eye margins, pale hairs; antenna dark; basoflagellomere oval, brownish centrally; bf = 1.2; ventral part of the frontal prominence brownish, with sparse pollinosity and with a tapering pollinose stripe extending from the eye margin to the antennae; black face, with long golden yellow hairs, except for a central, bare, shiny stripe extending from the bottom of the frontal prominence to the mouth edge; lower part of face with a white pollinose stripe extending from the eye margin to the mouth edge laterally; occiput white pollinose, with long hairs mostly golden yellow but, on the upper third of occiput, with a line of intermixed black hairs.

Thorax: Black; scutum with two medial, grey pollinose stripes extending for about the anterior three-quarters of the scutum; scutum mainly with golden yellow hairs, but scattered black hairs on the surface of the scutum posterior to TS; PAPT posterodorsally, NP, and PC with tufts of golden yellow hairs; dark-brown scutellum, black on lateral corners; scutellum with intermixed long pale hairs and scattered shorter black hairs; profemur black on the basal half, but black colour extending into the apical half on the posterior face of femora only; pro- and mesotibia and tarsi red; mesofemur black on the basal half; metafemur black on the basal two-thirds; metatibia darkened on the apical half; metatarsus red; all femora, tibiae, and tarsi with both pale and black hairs, but pro- and mesotarsus only with a few scattered black hairs; wings extensively microtrichose; wings light-brown pigmented on about the anterior third (cell R₂₊₃ very lightly pigmented).

Abdomen: Terga II–IV each with two lateral red markings covering the full length of the lateral margins of terga II and III (on tergum IV red markings are reduced to faint narrow bands on the anterior margin); tergum I with a moustache arrangement of golden yellow hairs with a central gap of about a third of the tergum width; terga II–IV with pale hairs except for the posterior margin, which has a band of black hairs of, at least, half of the tergum length.

Genitalia: Superior lobes about twice longer than wide, with an outwards-pointing process anteriorly (Fig. 58); surstyli, viewed in profile, with an anterior triangular expansion and two posterior excavations; surstyli with scattered long hairs posteriorly and apically (Fig. 59).

Head: Eye hairs brown, lighter and sparser lower down; vertical triangle shiny black, only pollinose on the anterior corner; ocellar triangle with long black hairs; frontal triangle shiny black, only pollinose on the top corner; frontal triangle only with long, anteriorly directed, black hairs; antenna black, slender, elongate, and thinly pollinose; scape conspicuously shorter than pedicel or basoflagellomere; basoflagellomere tapering towards the apex, as long as the pedicel or slightly longer (Fig. 51); bf = 3.8; arista red; profile of the face straight, or nearly so, between the antennae and the mouth edge; facial tubercle very slightly protruding; area just below the antennae black; face extensively shiny black with white pollinosity on the eye margins only (this marginal pollinosity is interrupted for a short distance on the genae, and occurs again on the eye margins); face with long white and black hairs intermixed, except for a central bare stripe between the antennae and the mouth edge.

Thorax: Scutum with two medial pollinose stripes extending slightly beyond the TS; scutum mostly with slightly posteriorly directed black hairs and, on the half of the scutum in front of the TS, scattered white hairs intermixed; PAPT posterodorsally, PP, NP, and AEP only with black hairs (NP and PAPT posterodorsally with higher density of hairs); anterior spiracle dark brown; posterior spiracle light brown; scutellum dark brown, black on the lateral corners; scutellum with long light-yellow and black hairs intermixed; legs extensively black, only red to dark red on the apex of metatibia and on the meso- and metatarsomeres 1–2 dorsally; legs with hairs mostly black, although with light-yellow hairs on the tibiae and tarsi, as well as on small areas of the metafemur basally; anterior margin of the wing conspicuously dark-brown pigmented (pigmented area tapering towards the apex); cells BM and CuP extensively bare of microtrichia; small areas of the cells DM, CuA1, and anal lobe bare of microtrichia; calypteres black.

Abdomen: Shiny black; tergum I with a moustache arrangement of light-yellow hairs; tergum II mostly with short black hairs, but also with scattered short light-yellow hairs intermixed; tergum II with long black hairs on the lateral margins, shorter when more posterior on the margins; terga III–IV with both very short light-yellow and black hairs intermixed; lateral margins of terga III–IV with hairs longer than those on the central plate; sterna mostly with long light-yellow hairs.

Genitalia: Superior lobes round at the apex, sinuous in shape; hypandrium with a conspicuously large hook-like process (Fig. 60); surstyli with round lateral expansions (Fig. 61).

Head: Eye hairs brown and denser on the upper third of the eye, with white, sparser hairs on the remaining area; ocellar triangle with black hairs; frons with black laterally directed hairs and sparse brown pollinosity medially on the eye margins from the frontal prominence to the ocellar triangle; dorsal area of the frontal prominence with anteriorly directed hairs; frons and frontal prominence dorsally with light-yellow hairs; scape and pedicel extensively black, only narrowly red on the pedicel apically; basoflagellomere oval (Fig. 12), dark red; bf = 1.4; ventral area of the frontal prominence white pollinose; face black, with long yellowish white hairs, except for a bare central shiny stripe from the bottom of the frontal prominence to the mouth edge; gena with a white pollinose stripe extending from the eye margin to the mouth edge; gena and occiput white pollinose on the eye margin; occiput with long yellowish white hairs and, on the upper third, a line of long black hairs intermixed.

Thorax: Black; scutum with two brownish pollinose approximated stripes extending beyond the TS but not reaching the posterior margin of the scutum; scutum with yellow hairs, but also with black hairs intermixed on the part of the scutum behind the TS; PAPT posterodorsally, NP, and PC with tufts of golden yellow hairs, not very dense on PC; scutellum dark brown, black on the lateral corners; scutellum with pale hairs only; all femora extensively black (pro- and mesofemur red on the apical half anteriorly); protibia red with darkened areas on the apical half; mesotibia red; metatibia red with darkened areas on about the central two thirds; tarsi red; femora and tibiae mostly with yellow hairs, but in all cases with some scattered black hairs or setae; wings wholly microtrichose, with the base of the anterior half lightly brown pigmented on the areas outlining the veins.

Abdomen: Tergum I with a moustache arrangement of laterally directed golden yellow hairs, separated by a gap shorter than a third of the tergum width; terga II extensively black, with two lateral triangular red markings extending on part of the lateral margins; tergum III with two lateral smaller red markings restricted on the anterior margin; terga II–IV posteriorly fringed with yellow hairs; terga II–IV with a band of black hairs on the posterior third of the tergum and reaching the lateral margins; tergum V mostly black haired.

Head: Eye hairs brown, but hairs sparser lower down; vertical triangle black with white pollinosity only on the anterior corner; ocellar triangle with long black hairs; frontal triangle black with long black hairs, but with light-red hairs on the eye margins; lunnules brownish; antenna extensively black, only with pedicel dark red apically and basoflagellomere dark red basally; basoflagellomere round; red arista, darkened apically; ventral area of the frontal prominence brownish, with sparse pollinosity; face black, with long brown to yellow hairs, except for a bare central shiny stripe from the bottom of the frontal prominence to the mouth edge; gena with a stripe of white pollinosity from the eye margin to the mouth edge; eye margins pollinose on the gena and the whole occiput; occiput with long white hairs and, on the upper third, scattered long black hairs intermixed.

Thorax: Scutum black pollinose, with two medial very faint greyish stripes not extending beyond the TS; scutum with yellow hairs on the half in front of the TS and black hairs on the half behind the TS; NP and PAPT posterodorsally with tufts of golden yellow hairs; AEP with long black hairs dorsally; PC with black hairs, but not arranged in tufts; anterior and posterior spiracles black; scutellum dark brown, even darker on the lateral corners; scutellum with black hairs centrally and long yellow hairs posteriorly; all femora extensively black, with red markings on about the apical fifth; pro- and mesotibia red; metatibia extensively dark brown, only red basally and apically; tarsomeres 1–2 red dorsally, but tarsomeres 3–5 black; all femora and tibiae with both yellow and black hairs; wings wholly microtrichose, with light-brown pigmentation anteriorly.

Abdomen: Tergum I with two triangular groups of golden yellow hairs obscuring the background colour of the tergum, separated for a distance of less than a quarter of the tergum width; terga II–IV red laterally, with red erect hairs but also with a semicircular band of posteriorly directed hairs on the posterior margin; sterna I–II black overall, but sterna III–IV black only centrally.

Genitalia: Superior lobes claw shaped (Fig. 62); anterior part of the surstyli gently excavated medially; surstyli with long hairs posteriorly (Figs 63, 64).

Head: All eye hairs white; ocellar triangle with both long, slightly anteriorly directed, black and white hairs; ocellar triangle with white pollinosity on the anterior corner; frontal triangle shiny black, with a hump on the upper third, conspicuous in lateral view; frontal triangle with long, anteriorly directed, pale hairs and, on the hump, a few black hairs; basoflagellomere rectangular, nearly 1.5 × as long as the pedicel, but conspicuously broader than this (Fig. 22); bf = 1.5; face black, with white pollinosity denser on the eye margin and on the gena; face with a central shiny stripe restricted to the facial tubercle, and not reaching the antennae; face with two lateral stripes thinly pollinose; face pale haired except for the central bare stripe; occiput pollinose.

Thorax: Scutum black, coarsely punctured, with two medial white pollinose stripes extending beyond the TS; scutum and pleuron pale haired; scutum with short, posteriorly directed hairs; NP, PAPT, AEP, dorsal katepisternum, and PC with hairs longer than those on the scutum; pleural plates without tufts of hairs; anterior and posterior spiracles dark brown; scutellum dark brown, with pale hairs; legs black, except for pale apex of the femora and base of tibiae; legs extensively pale haired, except for scattered short, black, adpressed hairs on the apical part of the metafemur dorsally; wings conspicuously brown pigmented on the anterior margin, although lightly in cell C; cells DM, CuP, CuA1, anal lobe, alula, and C with areas bare of microtrichia or more sparsely microtrichose than the rest of the wing.

Abdomen: With lateral margins nearly parallel; terga extensively black, pale haired; tergum I without a moustache arrangement of hairs; lateral margins of tergum I posteriorly and of tergum II anteriorly with long pale hairs, most of these conspicuously wavy in the apical third to half of the hair; sterna with long white hairs.

Genitalia: Superior lobes reduced (Fig. 65); surstyli straight, about bar-shaped (Fig. 66).

Head: Eye hairs straight and brown, slightly lighter and sparser downwards on the eye; vertical triangle black with white pollinosity on the anterior corner; ocellar triangle with long black hairs; frontal triangle black, with long black hairs, except for the eye margins bearing a line of pale hairs; basoflagellomere oval and brownish; bf = 1.4; ventral area of the frontal prominence brown with sparse pollinosity; face black, with long golden yellow hairs, except for a bare median shiny stripe from the bottom of the frontal prominence to the mouth edge; gena with a stripe of white pollinosity from the eye margin to the mouth edge; eye margins with pollinosity on gena and the whole occiput; occiput with long, golden yellow hairs and, on the upper third, a line of long black hairs.

Thorax: Black; scutum with two stripes of grey pollinosity extending on the anterior three-quarters of the scutum; scutum with golden yellow hairs and, on the half of the scutum behind the TS, a few black hairs intermixed; also black hairs on the area above the wing insertion; PAPT posterodorsally, NP, and PC with tufts of golden yellow hairs; scutellum dark brown, black on the lateral corners; scutellum mostly golden yellow haired; profemur black on the basal half and slightly darkened on the remaining area; pro- and mesotibia pale; pro- and mesotarsus pale; mesofemur darkened on the basal half; metafemur black on the basal two-thirds and lighter on the remaining area; metatibia dark brown; metatarsus pale, except for the darkened dorsal area of tarsomere 5; legs with both pale and black hairs on femora and tibiae (very few and localized black hairs on mesolegs); pro- and mesotarsus with only yellow hairs, although scattered dark-yellow hairs are intermixed; wings wholly microtrichose; wings dark-brown pigmented on the anterior third, especially on the whole cell R₂₊₃, except for the hyaline loop.

Abdomen: Terga II–IV each with two red markings, which, on tergum IV, are faint, narrow, and extend along most of the lateral margins; terga II-III red on the full length of the lateral margins; tergum I with a moustache arrangement of golden yellow hairs, with a central gap of about a third of the tergum width; terga II–IV pale haired, except for the black hairs on the posterior third of tergum II and posterior half of terga III–IV.

Genitalia: Superior lobes developed and curved forwards in the shape of a ‘reptilian’ head (Fig. 67); superior lobes with two lateral expansions (Fig. 67); surstyli heavily excavated anteriorly (Fig. 68).

Head: Eye hairs brown, except white and sparser lower down; vertical triangle shiny black, only pollinose on the anterior corner; ocellar triangle with long black hairs; frontal triangle shiny black, with long, anteriorly directed, black hairs; antenna thinly pollinose, black; arista red; basoflagellomere 1.5 × longer than the pedicel; scape conspicuously shorter than the basoflagellomere (Fig. 52); bf = 2.1; face straight between the bottom of the frontal prominence and the mouth edge, only the facial tubercle protruding; area just below the antennae narrowly yellow; face extensively shiny black, white pollinose on a narrow band along the eye margin (this band reaches the frontal prominence, above, and the gena, below, where it is interrupted for a short distance to continue along the eye margin up to the lower third of the occiput); face with long light-yellow hairs except for a central bare stripe extending from the antennae to the mouth edge.

Thorax: Scutum with two faint medial pollinose stripes extending beyond the TS; lateral pollinose stripes as well as the pollinose spots near the PP very faint, almost absent; scutum with slightly posteriorly directed red hairs, except for the black hairs on the lateral margins; PP with black setae; PAPT posterodorsally, NP, and AEP only with black hairs, denser on PAPT posterodorsally and NP; anterior spiracle dark brown; posterior spiracle light brown; dark-brown scutellum, black laterally and with long red hairs; legs extensively black, only metatibia apically and mesobasotarsomere red dorsally (metabasotarsomere dark red dorsally); legs mostly with black hairs, only with red hairs on tibiae, tarsi, and some reduced areas of the metafemur; wings heavily dark-brown pigmented on the anterior half (pigmented area tapering towards the wing apex) (Fig. 69); cells BM and CuP with lower density of microtrichia than other cells; calypter black.

Abdomen: Shiny black, mainly covered with short black hairs, except for the moustache arrangement of golden yellow hairs on tergum I; tergum II with long black hairs on the lateral margins, shorter when more posterior on the tergum; sterna with both long red and black hairs.

Genitalia: Shape of the superior lobes and surstyli as depicted in Figures 70–72.