Positive Periodic Solutions of Cooperative Systems with Delays and Feedback Controls
Abstract
This paper studies a class of periodic n species cooperative Lotka-Volterra systems with continuous time delays and feedback controls. Based on the continuation theorem of the coincidence degree theory developed by Gaines and Mawhin, some new sufficient conditions on the existence of positive periodic solutions are established.
1. Introduction
Mathematical ecological system has become one of the most important topics in the study of modern applied mathematics. Its dynamical behavior includes persistence, permanence and extinction of species, global stability of systems, the existence of positive periodic solutions, positive almost periodic solutions, and strictly positive solutions. The existence of positive periodic solutions has already become one of the most interesting subjects for scholars. In the recent years, the application of fixed-point theorems to the existence of positive periodic solutions in mathematical ecology has been studied extensively, for example, Brouwer’s fixed point theorem [1–4], Schauder’s fixed-point theorem [5–8], Krasnoselskii’s fixed-point theorem [9–14], Horn’s fixed-point theorem [15, 16], and Mawhins continuation theorem [17–36], and so forth. In particular, Mawhins continuation theorem is a powerful tool for studying the existence of periodic solutions of periodic high-dimensional time-delayed problems. When dealing with a time-delayed problem, it is very convenient and the result is relatively simple [30]. Recently, a considerable number of mathematical models with delays have been proposed in the study of population dynamics. One of the most celebrated models for population is the Lotka-Volterra system. Subsequently, a lot of the literature related to the study of the existence of positive periodic solutions for various Lotka-Volterra-type population dynamical systems with delays by using the method of continuation theorem was published and extensive research results were obtained [17–21, 24–34].
On the other hand, in some situations, people may wish to change the position of the existing periodic solution but to keep its stability. This is of significance in the control of ecology balance. One of the methods for the realization of it is to alter the system structurally by introducing some feedback control variables so as to get a population stabilizing at another periodic solution. The realization of the feedback control mechanism might be implemented by means of some biological control scheme or by harvesting procedure [21]. In fact, during the last decade, the existence of positive periodic solutions for the population dynamics with feedback control has been studied extensively [8, 14, 17–21, 24, 29]. To the best of our knowledge, studies on the existence of positive periodic solutions for cooperative systems with delays and feedback controls are fairly rare.
2. Preliminaries
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(H1) τijl(t) (l = 1,2, …, m, i, j = 1,2, …, n), σi(t), εi(t), ri(t) (i = 1,2, …n) are continuous ω-periodic functions with and . aijl(t) (i, j = 1,2, l = 1,2, …, m), βi(t), ei(t), bi(t), di(t), and γi(t) (i = 1,2, …n) are continuous, positive ω-periodic functions.
In order to obtain the existence of positive ω-periodic solutions of system (4), we will use the continuation theorem developed by Gaines and Mawhin in [36]. For the reader’s convenience, we will introduce the continuation theorem in the following.
Let X and Z be two normed vector spaces. Let L : Dom L ⊂ X → Z be a linear operator and let N : X → Z be a continuous operator. The operator L is called a Fredholm operator of index zero, if dim Ker L = codimIm L < ∞ and Im L is a closed set in Z. If L is a Fredholm operator of index zero, then there exist continuous projectors P : X → X and Q : Z → Z such that Im P = Ker L and Im L = Ker Q = Im (I − Q). It follows that L∣ Dom L ∩ Ker P: Dom L ∩ Ker P→ Im L is invertible and its inverse is denoted by KP; denote by J : Im Q → Ker L an isomorphism of Im Q onto Ker L. Let Ω be a bounded open subset of X; we say that the operator N is L-compact on , where denotes the closure of Ω in X, if is bounded and is compact.
Lemma 1 (see [35].)Suppose τ ∈ C1(R, R) with τ(t + ω) ≡ τ(t) and τ′(t) < 1, ∀t ∈ [0, ω]. Then the function t − τ(t) has a unique inverse function μ(t) satisfying μ ∈ C(R, R), μ(u + ω) = μ(u) + ω, ∀u ∈ R.
Lemma 2 (see [36].)Let L be a Fredholm operator of index zero and let N be L-compact on . If
- (a)
for each λ ∈ (0,1) and x ∈ ∂Ω ∩ Dom L, Lx ≠ λNx;
- (b)
for each x ∈ ∂Ω ∩ Ker L, QNx ≠ 0;
- (c)
deg {JQN, Ω ∩ Ker L, 0} ≠ 0,
3. Main Results
In order to obtain the existence of positive periodic solutions of system (4), firstly, we introduce the following lemma.
Lemma 3. Suppose that ,, , , is an ω-periodic solution of (4) and (5); then , , satisfies the system
Proof. By (4), (5), and the variation constants formula in ordinary differential equations, we have
Theorem 4. Suppose that assumption (H1) holds and there exists a constant θi > 0, ζi > 0, i = 1,2, …, n, such that
Proof. For system (18) we introduce new variables yi(t) (i = 1,2, …, n) such that
Now, we reach the position to search for an appropriate open bounded subset Ω for the application of the continuation theorem (Lemma 2) to system (26).
Corresponding to the operator equation Ly(t) = λNy(t) with parameter λ ∈ (0,1), we have
Assume that y(t) = (y1(t), y2(t), …, yn(t)) ∈ X is a solution of system (37) for some parameter λ ∈ (0,1). By integrating system (37) over the interval [0, ω], we obtain
From the assumption of Theorem 4, the equation has a unique positive solution . Hence, the equation QNy = 0 has a unique solution .
Choosing constant B > 0 large enough such that and B > B1 + B2 + ⋯+Bn, we define a bounded open set Ω ⊂ X as follows:
Acknowledgments
This work was supported by the Natural Science Foundation of Xinjiang (Grants no. 2012223177, 200721104) and the National Natural Science Foundation of China (Grants no. 10661010, 11261061).